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the difference is the .223 rem in a SAAMI minimum spec chamber, the wylde chamber is one that can utilize ammo that is either .223 remington or 5.56 Nato. The Nato and the Remington are close but slightly different. The .223 AI is an improved version of the .223 giving the shoulder a sharper angle and an advantage in case capacity and velocity.
On my DPMS AR-15 the gun is stamped 5.56. The guys at the gun shop stated that I can shoot 5.56 Nato and .223 Rem ammo caused the 5.56 has the higher chamber pressures of the two.
the wylde chamber will work with both the .223 and 5.56 but is designed to be a good medium . if you have a 5.56 chamber and shoot 223 chances are that it want be as acurate as if you had a 223 chamber. a 223 wylde gives you back some of that accuracy and still lets you shoot 5.56
i personally would buy a gun in the wylde chamber before the other 2.but 5.56 wold be my second choice