Yes, and no.
The AI wont go all the way up into the sliding sleeve. Just how much short kinda depends. You may or may not have enough adjustment in the seater stem to properly seat a bullet. Id recommend you call redding and see it they'll just sell you the AI sliding sleeve.
For the sake of making test dummies prior to a reamer order, i was able to seat a bullet in a .221 fireball case using a .223 die. This was on a Forster Br seater, which has a little more range in the seater stem than a redding. Not an everyday solution for sure.